Kileskus

Kileskus (meaning lizard in the Khakas language) is a genus of tyrannosauroid dinosaur known from partial remains found in Middle Jurassic (Bathonian stage) Itat Formation of Sharypovsky District, Krasnoyarsk Krai (Russia). Fossils recovered include the holotype maxilla, a premaxilla, a surangular, and a few bones from the hand and foot.[1] Additional remains referred to the species include cervical and caudal vertebrae, as well as a fibula.[2] The skull bones are similar to those of Proceratosaurus. The type species is K. aristotocus. Kileskus was named in 2010 by Averianov and colleagues. Its size has been estimated at 5.2 meters (17 ft) in length and 700 kg (1,540 lbs) in weight.[3]

Kileskus
Temporal range: Bathonian,
Diagram showing fossil remains described in initial description
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: †Proceratosauridae
Genus: †Kileskus
Averianov et al., 2010
Species:
†K. aristotocus
Binomial name
†Kileskus aristotocus
Averianov et al., 2010
Estimated size, compared to a human

Classification

Kileskus has been included in two phylogenetic analyses and found to be a basal proceratosaurid both times.[1][4]

Although it is unknown whether Kileskus sported a nasal crest, it can be assigned to Proceratosauridae due to a number of other features. These include elongated external nares, a short ventral margin of the premaxilla, and the area of the antorbital fossa directly below the antorbital fenestra being deeper than the maxilla directly below it. Kileskus also shares with Proceratosaurus nares inclined posterodorsally at a 40 degree angle to the skull. Kileskus is distinguished from other proceratosaurids by the anterior rim of its maxilla being confluent with the ascending process of the maxilla and gently sloping posterodorsally.[1]

Below is a cladogram published in 2013 by Loewen et al..[4]

Tyrannosauroidea
Proceratosauridae
 
 

Proceratosaurus bradleyi

 
 

Kileskus aristotocus

 
 

Guanlong wucaii

 
 
 
 

Sinotyrannus kazuoensis

 
 
 

Juratyrant langhami

 
 

Stokesosaurus clevelandi

 
 
 
 
 
 

Dilong paradoxus

 
 
 

Eotyrannus lengi

 
 
 

Bagaraatan ostromi

 
 
 

Raptorex kriegsteini

 
 
 

Dryptosaurus aquilunguis

 
 
 
 

Alectrosaurus olseni

 
 

Xiongguanlong baimoensis

 
 
 
 

Appalachiosaurus montgomeriensis

 
 
 
 

Alioramus altai

 
 

Alioramus remotus

 
 
 

Tyrannosauridae

 
 
 
 
 
 
 
 
 
 

Cladogram published in 2018.[5]

Tyrannosauroidea
Proceratosauridae
 
 

Guanlong wucaii

 
 
 

Proceratosaurus bradleyi

 
 
 

Kileskus aristotocus

 
 
 

Sinotyrannus kazuoensis

 
 

Yutyrannus huali

 
 
 
 
 
 
Pantyrannosauria
 
 

Aviatyrannis jurassica

 
 
 

Dilong paradoxus

 
 
 
 

Santanaraptor placidus

 
 

Timimus hermani

 
 
 
 

Stokesosaurus clevelandi

 
 

Juratyrant langhami

 
 

Eotyrannus lengi

 
 
 
 

Xiongguanlong baimoensis

 
 

NMV P186046 ("Australian tyrannosaur")

 
 
 
 

Alectrosaurus olseni

 
 

Timurlengia euotica

 
 
Eutyrannosauria
 

Dryptosaurus aquilunguis

 
 
 

Appalachiosaurus montgomeriensis

 
 
 

Bistahieversor sealeyi

 
 
Tyrannosauridae
Albertosaurinae
 

Gorgosaurus libratus

 
 

Albertosaurus sarcophagus

 
 
Tyrannosaurinae
Alioramini
 

Qianzhousaurus sinensis

 
 
 

Alioramus remotus

 
 

Alioramus altai

 
 
 
 
 

Nanuqsaurus hoglundi

 
 

Teratophoneus curriei

 
 

Lythronax argestes

 
 
 
 
 

Daspletosaurus torosus

 
 

Daspletosaurus horneri

 
 
 
 
 

Zhuchengtyrannus magnus

 
 
 

Tarbosaurus bataar

 
 

Tyrannosaurus rex

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 

See also

References

  1. Averianov, A. O.; Krasnolutskii, S. A.; Ivantsov, S. V. (2010). "A new basal coelurosaur (Dinosauria: Theropoda) from the Middle Jurassic of Siberia". Proceedings of the Zoological Institute. 314 (1): 42–57.
  2. O. Averianov, Alexander; Osochnikova, Anastasia; Skutschas, Pavel; Krasnolutskii, Sergei; Schellhorn, Rico; A. Schultz, Julia; Martin, Thomas (2019-09-16). "New data on the tyrannosauroid dinosaur Kileskus from the Middle Jurassic of Siberia, Russia". Historical Biology. 33 (7): 897–903. doi:10.1080/08912963.2019.1666839. ISSN 0891-2963. S2CID 203890300.
  3. Molina-Pérez & Larramendi (2016). Récords y curiosidades de los dinosaurios Terópodos y otros dinosauromorfos. Spain: Larousse. p. 264.
  4. Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C (ed.). "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE. 8 (11): e79420. Bibcode:2013PLoSO...879420L. doi:10.1371/journal.pone.0079420. PMC 3819173. PMID 24223179.
  5. Delcourt, R.; Grillo, O. N. (2018). "Tyrannosauroids from the Southern Hemisphere: Implications for biogeography, evolution, and taxonomy". Palaeogeography, Palaeoclimatology, Palaeoecology. 511: 379–387


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